I'll be darned..looks to me to be SLIGHT Grover Dents..when pic is blown up, I swear I see the ring around the Kluson bushing:
So TOTALLY possible these could be the original keys, wisely saved when the Grovers were installed..
then when re-installed, would be much cleaner and not match the age of the guitar..they are damn clean.
Would love to see two things, the peg head hole with the tuner removed, (to see if reamed), and a tuner close up, and the underside of one would confirm if repro or not..
When looking at them online, it's too easy to get burned, so I always ask to see the underside of the tuners...to see the PAT# of the Kluson.
If it had the PAT#, then damn, those are almost new looking, so if not repros, era correct replacements..OR new double ring keys put on is another possibility.